Joan Roughgarden

Joan Roughgarden is an American ecologist and evolutionary biologist. She has engaged in theory and observation of coevolution and competition in Anolis lizards of the Caribbean, and recruitment limitation in the rocky intertidal zones of California and Oregon. She has more recently become known for her rejection of sexual selection, her theistic evolutionism, and her work on holobiont evolution.

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  Second Nature: Gender and Sexuality in the Animal World

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• 2026Second Nature: Gender and Sexuality in the Animal Worldself